Article 2375 Carolina Royo Pablo Carbonell-Bejeran Rafael Torres-Pérez Anna Nebish Óscar Martínez Manuel Rey Rouben Aroutiounian Javier Ibáñez José M. Martínez-Zapater Journal of Experimental Botany 67 (1) 259-273 2016 http://jxb.oxfordjournals.org/